For the past two decades, Kay E. Holekamp has been chronicling the lives of spotted hyenas on the savannas of southern Kenya. She has watched cubs emerge from their dens and take their place in the hyena hierarchy; she has seen alliances form and collapse. She has observed clan wars, in which dozens of hyenas have joined together to defend their hunting grounds against invaders.

“It’s like following a soap opera,” said Dr. Holekamp, a professor at Michigan State University.