Two animal handlers who say they were fired for refusing to expose their breasts to a 300-pound gorilla have settled their lawsuit against the Gorilla Foundation on undisclosed terms.

Nancy Alperin and Kendra Keller, both in their mid-40s, claimed that Gorilla Foundation president Francine “Penny” Patterson pressured them to indulge Koko the gorilla’s “nipple fetish” as a way of bonding with the 33-year-old female.

The foundation denied the allegations.